1986-1990 Trinity College, University of Oxford. BA (1^st Class Honours) Biochemistry (MA 1993)
1990-1993 Trinity College, University of Oxford.  DPhil. Biochemistry.  

Current positions:

Since 2015  Professor of Biophysics, Department of Physics, Oxford .
Since 2008  Fellow, Green Templeton College

Previous positions:

1994-1995  Wellcome Trust International Prize Travelling Fellow. Vollum Institute, Portland, Oregon USA.
1996-2000  Wellcome Trust Career Development Fellow, University Laboratory of Physiology, Oxford.
1996-2008  RJP Williams Research Fellow, Wadham College , Oxford.
1999-2007  University Research Lecturer, Department of Physiology Anatomy and Genetics, Oxford
2001-2008  Royal Society University Research Fellow,  Dept Physiology Anatomy and Genetics, Oxford
2007-2014  University Lecturer in Biological Physics, Department of Physics, Oxford
2014-2015  Associate Professor of Biological Physics, Department of Physics, Oxford .